Table 2

Details of defects associated with malolactic fermentation (MLF).

DefectConditions increasing susceptibilityCompounds modifiedCompounds producedSensory effectsSource
Increased pHHigh pH, high climatic temperatures, uncontrolled/spontaneous MLF, malic acid additionMalic acidLactic acid; ethyl lactate and ethyl acetate; propanoic acid; 2-methyl; 3-hydroxy-2,2,4-trimethylpentyl esterFlavors described as lactic, wet wool/wet dog, rancid, cheesy, sweatyKeng et al. 2025
Color instabilityHigh pH, high climatic temperatures, oxygen exposureAnthocyaninsGlycosidases, anthocyanidins, monomeric anthocyaninsReduced color intensity, oxidation with increased pHTofalo et al. 2021, Burns and Osborne 2013, Bartowsky and Krieger-Weber 2020
Increased alcohol/fusel alcoholsHigh pH, high climatic temperatures, high alcohol, oxygen exposurea-Keto acidsFusel aldehydes, fusel alcoholsBurning/tingling sensation; flavors described as solvent-like, boozy, and bitterHazelwood et al. 2008, Lerm et al. 2010, de-la-Fuente-Blanco et al. 2024
RopinessHigh pH, high climatic temperatures, more common in white wineGlucoseExtracellular polysaccharides, glucanIncreased viscosity, gelatinous substancesCiezack et al. 2010, Dimopoulou and Dols-Lafargue 2021
MousinessHigh pH, oxidative conditions, uncontrolled/spontaneous MLFAmino acidsN-heterocyclic compounds, 2-acetyltetrahydropyridine, 2-acetyl-1-pyrrolineMousinessCostello et al. 2001
Citric acid degradationHigh pH, uncontrolled MLFCitric acidAcetic acid, diacetyl, acetoinVolatile acidity, buttery and nutty flavorsNielsen and Richelieu 1999
Tartaric acid degradationHigh pHTartaric acidLactic acid, acetic acidIncreased pH, volatile acidity, effervescence, lactic flavorsKrumperman and Vaughn 1966
Glycerol degradationHigh pH (>4.0), high climatic temperatures, high postharvest temperatures (>20°C), low alcohol/low fermentable sugar, press wines, wines with prolonged lees agingGlycerol3-Hydroxypropionaldehyde, acrolein, acetic acidBitterness, volatile acidityBauer et al. 2010, Popescu-Mitroi et al. 2014